Dr. Alicia Ely Joson-Pagulayan did a paper in UP in 1999 on the aspects of tawilis reproductive biology. Photo by Jo. Lontoc, UP MPRO
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Much has changed in UP over the past 100 years. From one campus in Manila, it now has eight constituent universities with 17 campuses all over the country; it has 258 undergraduate programs; and 438 graduate programs with students from almost every region in the country. Learn More
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